Historical biogeography of Cymbiodyta. A dated phylogeny of Cymbiodyta adopted from Toussaint and Short (2019), with C. samueli sp. nov. added as a sister species to modern C. marginella, with four alternative ages of their MRCA used for biogeographic analyses. B historical biogeography of Cymbiodyta following the DIVALIKE model in BioGeoBEARS (dotted rectangles show nodes for which the ancestral reconstruction diferred in DIVALIKE+j analysis). C, positions of the continents at the end of the Cretaceous (Albian) when European and American Cymbiodyta separated (NA, North America; SA, South America; E, Europe; AF, Africa; AS, Asia).