Biogeographic hypothesis of the Wormaldia occipitalis species complex. A–D Biogeographic dynamics of the (sub)groups along time. E Recent distribution of the (sub)groups of the W. occipitalis species complex. Individuals are colour-coded by their morphological determination. Blue – W. occipitalis occipitalis, green – W. occipitalis meridionalis, gold – W. occipitalis x subterranea, red – W. subterranea, Grey – W. sp. Genetic groups were also colour-coded and additionally labelled. Light blue – W. occipitalis ssp., pink – W. subterranea ssp.