Research Article
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Article title
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Keywords
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1. Introduction
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2. Material and methods
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2.1. Material studied
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2.2. Microscopy and image processing
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2.3. Dataset and morphological treatment
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2.4. Cladistic analysis
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2.5. Nomenclatural act
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3. Results
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3.1. Morphological characters list
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3.2. Morphological phylogeny
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3.3. Family Gripopterygidae and Subfamily Arrangements
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3.4. Subfamily Gripopteryginae
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3.5. Systematics
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Tupiperla
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Tupiperla furcata
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4. Discussion
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4.1. Redefinition of Gripopteryginae
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4.2. Gripopteryginae (sensu stricto) and related taxa
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4.3. The Case of Paragripopteryx munoai
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4.4. Polyphyly in Leptoperlinae and Dinotoperlinae
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5. Conclusions and Future Directions
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6. Declarations
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7. Acknowledgements
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8. References
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Supplementary materials
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