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  • ContentsContents
  • Article InfoArticle Info
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  • Article title
  • Abstract
  • Key words
  • 1. Introduction
  • 2. Material and methods
    • 2.1. Sample collection, identification and DNA extraction
    • 2.2. Mitogenome sequencing and assembly
    • 2.3. Mitogenome annotation and analysis
    • 2.4. Phylogenetic analysis
    • 2.5. Divergence time estimation
    • 2.6. Ancestral area reconstruction
    • 2.7. Ancestral reconstruction of morphology
  • 3. Results
    • 3.1. Genome composition and base content
    • 3.2. Evolution of PCGs
    • 3.3. Phylogenetic relationship
    • 3.4. Biogeography of Hexarthrius
    • 3.5. Divergence time estimation
  • 4. Discussion
    • 4.1. Implications of gene rearrangements
    • 4.2. Biogeographical history of Hexarthrius
    • 4.3. Morphological evolution of Hexarthrius
    • 4.4. Species delimitation and cryptic species
  • 5. Declarations
  • 6. Acknowledgements
  • 7. References
  • Supplementary materials
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