New insights into the phylogeny of Tortopus Needham and Murphy and Tortopsis Molineri (Ephemeroptera, Polymitarcyidae) with description of three new species

Tortopsis toro sp. nov., nymph. 7, labrum, dorsal view; 8, left mandible, inner view, 9, right mandible, inner view; 10, right mandible, detail of tusk, ventral view (arrow indicates stout spine near inner margin); 11, hypopharynx, dorsal view; 12, labium, ventral view; 13, left maxilla, ventral view (arrow indicates membranous gill); 14, right maxilla, ventral view.