New insights into the phylogeny of Tortopus Needham and Murphy and Tortopsis Molineri (Ephemeroptera, Polymitarcyidae) with description of three new species

Tortopsis toro sp. nov., imago: 19, male genitalia, ventral view; 20, detail of apex of penis, ventrolateral view; 21, male genitalia, lateral view; 22, female abdominal sternum VIII. Nymph: 23, head outline and setation, dorsal view (tusks omitted); 24, left mandible, inner view (arrow indicates stout spine near inner margin); 25, detail of right mandibular tusk, inner view (arrow indicates subapical outer indentation); 26, paraprocts, ventral view (right half setation omitted).