New insights into the phylogeny of Tortopus Needham and Murphy and Tortopsis Molineri (Ephemeroptera, Polymitarcyidae) with description of three new species

Tortopus coreguaje sp. nov., male imago. 56, genitalia, ventral view (left forceps omitted); 57, genitalia, lateral view. Nymph: 58, head outline and setation, dorsal view (tusks omitted); 59, right mandible, inner view; 60, left mandible, inner view; 61, same, detail of apex (arrow indicates blade-like projection); 62, fore leg, ventral view (rows of filtering setae delineated); 63, fore tarsal claw; 64, hind tarsal claw.