Sertaneja gen. nov. resembles Gyndoides because of the ocularium armature (despite being variable in Sertaneja), four scute areas, a pair of paramedian spines on scute area II (except S. falcata sp. nov.) and unarmed free tergites. Sertaneja gen. nov. differs from Gyndoides Mello-Leitão, 1927a by: (i) the lateral margin of DS with an external row of tubercles slightly increasing in size posteriorly, (ii) presence of the mesal apical seta on the pedipalp femur (Figs 3F, 6F, 9F), (iii) male trochanter IV retro-apical projection as a straight apophysis instead of a tubercle (Figs 3D, 6D, 9D), (iv) the absence of a retro-basal apophysis on male femur IV, (v) male femur IV bearing a prodorsal apical distinguished apophysis, (vi) male tibia IV ventral face armed, (vii) glans without ventral process (Figs 5, 8, 11), (viii) stylus apex bearing distinct projections (Figs 5, 8, 11), (ix) VP of penis without prominent ventral lobes (Figs 5, 8, 11), and, (x) 4 pairs of MS A clustered instead of lined up in a row (Figs 5, 8, 11). It differs from other Pachylinae genera by the combination of the sub-rectangular VP, 2 paralateral pairs of MS D, glans stylus bearing a dorso-apical longitudinal projection and lack of ventral process of the glans.

DS gamma (γ) to alpha (α) shaped (Figs 3A, B, 6A, B, 9A, B). Ocularium height at least two to up to three times the size the eye diameter, with posterior face tuberculate, and slightly close to the anterior margin of DS, in lateral view (Figs 3C, 6C, 9C); bearing a pair of divergent large spines (Figs 3G, 9G), or a single central robust one (Fig. 6G). Four scute areas; scute areas I, II and IV with one to two pairs of slightly enlarged paramedian tubercles; scute area III with a pair of distinct paramedian elevations, ranging from acuminated tubercles (Fig. 9A, C) to large posteriorly curved spines (Figs 3A, C, 6A, C). Chelicerae segment I posterior face covered with acuminated tubercles. Pedipalp femora with a mesal sub-apical setiferous tubercle. Coxa IV well developed, bearing dense and high tuberculation, with robust prodorsal apical apophysis. Trochanter III with a retro-ventral apical conical tubercle. Trochanter IV with a straight retro-apical conical apophysis (Figs 4A, C, D, 7A, C, D, 10A, C, D), and a short probasal apophysis. Penis VP sub-rectangular (Figs 5A, 8A, 11A); glans without dorsal or ventral processes (Figs 5B, 8B, 11B), stylus with a dorso-apical projection slightly curved posteriorly, ventral face with sub-apical transversal row of trichome-like projections. MS A group generally composed of four to five pairs of paralateral setae, at instances showing asymmetry between the sides (Fig. 8A). MS C composed by three (Figs 5, 8) or five pairs of setae (Fig. 12) inserted laterally on the apical third of the VP. MS D as two pairs of paralateral setae, inserted more dorsally on the VP, between de MS A and C sets. MS E as a pair of paralateral small setae, inserted ventrally near the MS C and D groups. Ventral surface of VP sparsely covered by microsetae.

S. bicupidata sp. nov., S. crassitibialis sp. nov., and S. falcata sp. nov.

S. bicuspidata sp. nov.

‘Sertaneja’ is a Brazilian adjective (fem.) that refers to a woman who lives in the Sertão regions in rural communities often in harsh survival conditions. Sertão is the largest sub-region of Brazilian Northeast characterized by dry climates and Caatinga vegetation.

BRAZIL, Ceará: Ubajara, Parque Nacional de Ubajara.

Holotype. BRAZIL. Ceará: Ubajara PARNA de Ubajara, Rio Gameleira, elev. 840 m, 27.i.2014, DaSilva M.B. Saraiva N.E.V. and Sampaio C., ♂ (UFPB OP-163) [dissected]. Paratypes. BRAZIL. Ceará: Quixadá, Serra do Estevão, estrada para Dom Maurício, elev. 482 m, 18.iv.2014, DaSilva M.B. and Saraiva N.E.V., 1 ♂ (UFPB OP-724) [dissected]; Quixadá, Serra do Urucu, Santuário de Nossa Senhora Imaculada Rainha do Sertão, elev. 344 m, 19.iv.2014, DaSilva M.B. and Saraiva N.E.V., 1 ♂ 2 imm (MZUSP 76645) [male genitalia used for SEM], 1 ♂ (MNRJ 60616) [dissected]; Quixadá, Jardim dos Monólitos, inselbergs do Açude Euclides, elev. 180–200 m, 20.iv.2014, DaSilva M.B. and Saraiva N.E.V., 1 ♀ (UFPB OP-726) [female description].

This species can be distinguished from the other species of the genus by the denser tuberculation of the body surface (Fig. 3A); scute area III paramedian pair of spines enlarged and slightly more robust when compared to S. crassitibialis sp. nov. (Fig. 3A, C); coxa IV with uniquely shaped bifid prodorsal apical apophysis and conical retro-apical apophysis (Fig. 3A, D); trochanter IV with the largest retro-apical apophysis among the genus, and a large proapical apophysis curved dorsally (Figs 3D, 4); femur IV with a retro-lateral row of large conical apophyses (Fig. 4A, D). It most resembles S. falcata sp. nov. by the male femur IV dorsal row with equally sized high tubercles; penis glans’ stylus straight and with a rounded dorso-apical projection; VP macrosetae conical, MS A and MS B sets adjacent, and MS D pairs closer to each other and to MS C set (Fig. 5).

Holotype MALE (UFPB OP-163; Figs 3–4): Measurements: DSL: 4.86; DSW: 4.97; LI: 9.13; LII: 17.62; LIII: 10.78 (broken metatarsus); LIV: 15.48. Dorsum (Fig. 3A, C, G): DS anterior margin with a pair of small tubercles on each side. Prosoma covered with distinctive large tubercles concentrated on the middle of posterior half. Ocularium tuberculate, with a pair of spines slightly longer than the ocularium height, and a pair of large tubercles just behind (Fig. 3C). Two pairs of ozopores of similar size, anterior one placed more laterally, and posterior one more rounded and slightly dislocated dorsally; lateral channel well developed followed by distinguished tuberculate oblong sensorial pegs. Scute areas I–IV densely tuberculate, tubercles increasing in size medially. Scute area III paramedian pair of spines recurved, slightly reaching scute area IV. Lateral margin of DS with an external row of tubercles increasing in size towards scute area II, becoming slightly acuminated (Fig. 3C). Posterior margin of DS and free tergites I–III each with a transversal row of tubercles. Free tergite I with tubercles increasing in size medially. Anal operculum tuberculate. Venter: Coxa I with three longitudinal rows of tubercles increasing in size apically, median row largest; coxae II–III with slightly enlarged apical tubercles; coxa IV tuberculate. Chelicerae: Segment I with a transversal row of five tubercles on the posterior face of bulla. Fixed and movable fingers with five and four teeth; respectively. Pedipalps (Fig. 3E, F): Trochanter inflated dorsally, ventral face with a pair of setiferous tubercles, mesal largest (Fig. 3E). Femur with a dorsal median row of five tubercles, ventral face with a basal large setiferous tubercle (Fig. 3E, F); an external lateral row of five tubercles. Tibial setation: mesal and lateral IiIi/Iii. Tarsal setation: mesal IIi, lateral IiIi. Legs (Figs 3D, 4): Coxae I–IV each with one prodorsal and one retro-dorsal apophysis; coxa II with a trifid retro-lateral apophysis. Coxa IV bearing a robust, bifid, prodorsal apical apophysis, almost square-shaped, the large branch pointing backwards and the smaller one almost transversal, obliquely angled (Fig. 3D); and a conical retro-lateral apophysis, comparable in size and shape with the small, transversal branch of the prolateral one (Fig. 3D). Trochanter IV apically swollen in dorsal view, prolateral surface with a short, blunt, conical, basal apophysis, and a large, apical apophysis, perpendicular in lateral view, dorsally curved along its entire length, bearing a basal small acute straight apophysis pointing posteriorly (Fig. 4B); retro-lateral face with a basal short conical apophysis (ca. same size as the coxa IV retro-lateral apophysis), a sub-apical small, straight, transversal conical apophysis, and a long, straight apical apophysis 2/3 the length as the podomere (Fig. 4). Femur III slightly curved in dorsal view, with a paralateral pair of dorso-apical tubercles (retro-dorsal largest); ventral surface with two rows of tubercles increasing in size apically from their apical half, prolateral ones larger. Femur IV slightly curved inwards and upwards; dorsal surface with a median row of five enlarged, blunt, conical tubercles placed between the sub-basal and sub-apical regions of the podomere (Fig. 4B); dorso-apical surface with two paralateral large spines; prolateral row of low rounded tubercles; retro-lateral row of seven high apophyses roughly decreasing in size apically, the large basal ones approximately the same size as the trochanter IV retro-apical apophysis (Fig. 4A); two ventral rows of tubercles, proventral row with seven enlarged, blunt, conical tubercles on the basal half, retro-ventral row of small rounded tubercles (Fig. 4C); ventro-apical surface with a pair of paralateral enlarged acuminated tubercles, ca. three times the size of the immediately following tubercles (prolateral largest). Patella IV covered with conical tubercles that increase in size apically, ventral face with two rows of tubercles increasing in size apically, retro-lateral ones larger. Tibia IV covered by conical tubercles, ventral row of tubercles slightly increasing in size apically (Fig. 4B, D). Tarsal counts: 6, 10, 6, 6. Penis (UFPB OP-725; Fig. 5): Glans stylus cylindrical. VP apex convex with a slight median concavity, lateral margins slightly concave presenting a subtle constriction on its medial portion. VP bearing three paralateral pairs of MS A in a tight cluster in lateral view (Fig. 5B); one pair of MS B of same size as MS A, placed more basally and adjacent to the MS A cluster; three pairs of MS C, the apical spine slightly smaller; MS D set closer to MS C group, setae close to each other; and two pairs of reduced, sub-apical, ventro-lateral MS E positioned between the MS C setae (not visible in the SEM photograph, but confirmed in the other dissected males). Coloration: DS, tergites I–III, chelicerae, pedipalps coxae, coxae I–IV, and leg IV general coloration strong yellowish brown (74). DS with posterior and lateral margins bearing a moderate yellowish brown (77) outline; scute areas, prosoma and ocularium with moderate yellowish brown (77) granulation. Pedipalps trochanter–tibia, and legs I–III dark yellow (88).

FEMALE (UFPB OP-726; Fig. 3B): Measurements: DSL: 4.34; DSW: 4.20; LI: 8.32; LII: 15.48; LIII: 10.77; LIV: 14.27. Dorsum: DS gamma (γ), but coda section is slightly longer. Ocularium spines length as tall as the ocularium. Scute area III central armature shorter than male and more acuminated. Scute area IV with median pair of tubercles more conspicuous than male. Posterior margin of DS and free tergites I–III each with leaner tubercles on the sides. Legs: Coxa IV narrower and shorter than in male, with a short, conical, proapical apophysis obliquely inserted; retro-apical apophysis reduced. Trochanter III retro-apical projection smaller than in male. Trochanter IV shorter than in male, apical half swollen, bearing a terminal tubercle, with unarmed prolateral face; retro-lateral face with short basal apophysis, and blunt apical apophysis smaller than in male. Femur III ventral row and ventro-apical tubercles smaller than in male. Femur IV dorso-medial row with small and slender projections on basal 4/5; a retro-lateral, a proventral and a retro-ventral rows of small conical tubercles; dorso-apical and ventro-apical paralateral projections reduced. Tibia III proventral and retro-ventral rows of small granule-like tubercles. Patella–tibia IV dorsal face covered by conical blunt tubercles. Tarsal count: 6, 9–10, 6, 6.

Males (n = 4): Measurements: DSL: 4.33–4.86; DSW: 4.47–4.97; LI: 8.52–9.13; LII: 15.60–17.62; LIII: 10.78–12.30; LIV: 14.25–15.83. Dorsum: Ocularium spines as tall as, or slightly taller than the ocularium. Scute area III paramedian pair of spines medium to large, length reaching backwards up to halfway of scute area IV. Pedipalps: Pedipalpal setation: tibial mesal IiIi, lateral IiIi/i-Ii; tarsal mesal IIi/IIii, lateral IiIi/Iii. Legs: Minor males, compared to major males as the holotype, with less developed armature on scute area III and weaker armature of legs: coxa IV narrower and shorter, prodorsal apical and retro-apical apophyses smaller and slender (on major males prodorsal apical apophysis shorter projection can be acuminate); trochanter III retro-lateral projection reduced; trochanter IV all apophyses smaller; femur III prolateral and ventral rows of tubercles less developed, with smaller proapical tubercles; femur IV pro- and ventro-apical tubercles smaller; tibia IV proventral and retro-ventral rows reduced. Tarsal count: 6, 9–10, 6, 6.

‘Bicuspidata’ is a Latin adjective (adj. perf. part., nom. fem.) that refers to the diagnostic bifid apophysis of male coxa IV in this species.

Caatinga shrublands and dry forests of northern interior Ceará and Brejos de Altitude humid montane forests of Ibiapaba mountains.

BRAZIL, Ceará: Quixadá, Serra do Urucu, Santuário Nossa Senhora Imaculada Rainha do Sertão.

Holotype. BRAZIL. Ceará: Quixadá, Serra do Urucu, Santuário Nossa Senhora Imaculada Rainha do Sertão, elev. 344 m, 19.iv.2014, DaSilva M.B. and Saraiva N.E.V., ♂ (UFPB OP-487). Paratypes. BRAZIL. Ceará: Ubajara, PARNA de Ubajara, Brejo de Altitude, 700 m, 27.i.2014, DaSilva M.B., Saraiva N.E.V. and Sampaio C., 1 ♂ (MNRJ 60614) [genitalia used for SEM]; Ubajara, PARNA de Ubajara, Rio Gameleira, 840 m, 27.i.2014, DaSilva M.B.; Saraiva N.E.V. and Sampaio C., 2 ♀ (UFPB OP-480), 1 ♀ (UFPB OP-727) [female description]; Ubajara, PARNA de Ubajara, Gr. Morcego Branco, Gr. Ubajara, Trilha Portão Planalto, 22.x.2011, DeSouza A.M., Araujo E.S., Carvalho L., 1 ♀ (MNRJ 60615); Ubajara, PARNA de Ubajara, Trilha da Samambaia, 30.iv.2018, Silvino A.C.S. and Saraiva N.E.V., 1 ♂ 1 ♀ 1 imm. (MZUSP 76646) [male dissected].

Differs from the other species by the lower density of tubercles and armature (Fig. 6A); ocularium is the tallest in the genus and bears a robust, single spine (Fig. 6C); lateral margin of DS with a large acuminate tubercle on the external row, placed close to scute area III (Fig. 6C); scute area III paramedian pair of spines slender when compared to S. bicuspidata sp. nov. (Fig. 6C); coxa IV with an unbranched, robust prodorsal apical apophysis, missing a retro-apical apophysis (Fig. 6D); trochanter IV short, square-shaped with a robust, blunt central apophysis (Figs 6D, 7); femur IV less armed; tibia IV sexually dimorphic, swollen in males (Fig. 7); penis glans’ stylus with an angular dorso-apical projection (Fig. 8B); VP macrosetae spatulated, except for the basal MS D pair and MS E; MS B inserted very basally; and MS D pairs much apart from each other (Fig. 8).

Holotype MALE (UFPB OP-487; Figs 6, 7): Measurements: DSL: 4.91; DSW: 4.67; LI: 9.35; LII: 16.17; LIII: 12.44; LIV: 16.14. Dorsum (Fig. 6A, C, G): Prosoma sparsely covered by small granule-like tubercles. Ocularium twice as tall as the eye diameter, with a robust, conical spine projected anteriorly, slightly larger than the ocularium (Fig. 6G). Two pair of ozopores placed on the lateral surface, both slit shaped; lateral channel well developed, conspicuous and with small sensorial pegs. Scute area II, III and IV with a central transversal row of tubercles (Fig. 6A). Scute area III paramedian pair of spines slightly curved posteriorly in all its extension, barely reaching scute area IV (Fig. 6A). Lateral margin of DS with an external row of tubercles increasing in size posteriorly, becoming acuminated, largest one placed near scute area III (Fig. 6C). Posterior margin of DS and free tergites I–III each with a row of similar sized round tubercles. Anal operculum tuberculate. Venter: Coxae I–IV with low tubercles; coxa I with median row of enlarged tubercles. Coxa IV covered by granule-like tubercles. Chelicerae: Segment I with the posterior portion of the bulla bearing a transversal row of four tubercles. Fixed and movable fingers each with five and five to six teeth, respectively. Pedipalps (Fig. 6E, F): Trochanter inflated dorsally; ventral face with two setiferous tubercles, mesal tubercle largest. Femur with a dorsal median row of six tubercles (Fig. 6E); ventral surface with a large basal setiferous tubercle and a retro-lateral row of three tubercles on apical half. Tibial setation: mesal and lateral IiIi. Tarsal setation: mesal IIi, lateral IiIi. Legs (Figs 6D, 7): Coxae I–III each with one prodorsal and 1 retro-dorsal apophysis; coxa II retro-dorsal apophysis trifid and fused in the apex with coxa III prodorsal one. Coxa IV large, tuberculate, bearing a robust, unbranched prodorsal apical apophysis, this almost transversally inserted, slightly curved downwards and posteriorly (Fig. 6A, C, D). Trochanter IV with a robust, slightly bifid, blunt central apophysis, a prodorsal sub-apical enlarged tubercle, and an acuminated, straight retro-apical apophysis (Figs 6D, 7). Femur III with an enlarged retro-apical dorsal spine; ventral face with a pro- and retro-lateral rows of tubercles increasing in size apically on apical half. Tibia III sub-apical surface slightly swollen, with retro-ventral sub-apical row of enlarged tubercles. Femur IV slightly curved inwards and upwards; dorsal surface covered by small tubercles, with a pair of paralateral apical spines slightly curved upwards (prolateral spine largest) (Fig. 7A); lateral rows of tubercles slightly increasing in size apically, retro-lateral row largest; with pro and retro-ventral rows that increase in size apically, proventral row largest and with tubercles more close to each other (Fig. 7C); apically with a pair of paralateral spines, prolateral largest (Fig. 7C). Patella IV tuberculate, with a slightly swollen retro-dorsal apical spine (Fig. 7A). Tibia IV swollen at apical 3/4 of its length, with a proventral and retro-ventral row of enlarged sub-apical tubercles, and a pair of paralateral ventro-apical spines (Fig. 7B, C, D). Tarsal counts: 6, 8, 6, 6. Penis (UFPB OP-171; Fig. 8): VP apex slightly concave, lateral margin sub-straight. Proximal portion of VP with four pairs MS A and one pair MS B placed far from A4 and placed more basally. MS C set apically curved and showing asymmetry between left (3) and right (2) sides (Fig. 8A). MS D apical pair flattened and acuminate, basal spine conical and blunt; and two pairs of MS E placed more ventrally and close to MS C and MS D (Fig. 8B). Coloration: DS predominantly dark grayish yellowish brown (81) with scute areas limits, area III paramedian spines, and specks on prosoma dark orange yellow (72). Chelicerae, pedipalps, coxae I–III, and legs I–III dark yellow (88). Leg IV dark grayish yellowish brown (81).

FEMALE (UFPB OP-727; Fig. 6B): Measurements: DSL: 4.70; DSW: 4.26; LI: 8.88; LII: 15.36; LIII: 11.19; LIV: 14.81. Dorsum: Overall less developed armature and tubercles. DS alpha (α), coda slightly longer. Ocularium spine slender. Scute area III central armature small. Lateral margin of DS with external row of smaller tubercles than in male. Legs: Coxa IV narrower and shorter than in male, with an obliquely inserted, slender, conical, prodorsal apical apophysis, slightly curved downwards. Trochanter III retro-apical apophysis reduced. Trochanter IV narrower than in male, prolateral face unarmed, retro-lateral face as in males but with smaller apical apophysis. Femur III with ventral rows of small tubercles. Femur IV less armed, ventral rows of tubercles slightly increasing in size apically, dorso- and ventro-apical pair of spines small. Patella IV prodorsal apical projection reduced. Tibiae III–IV unarmed, not inflated. Tarsal counts: 6, 7, 6, 6.

Males (n = 3): Measurements: DSL: 4.91–4.99; DSW: 4.67–4.90; LI: 8.98–10.08; LII: 15.31–16.17; LIII: 11.58–12.44; LIV: 16.14–16.34. Dorsum: Ocularium spine more, or less acute. DS lateral margin external row of rounded to acuminate tubercles, becoming almost spine-like. Scute area II with tubercles placed near the lateral margins. Legs: Tibia IV retro-ventral apical spine single or bifid. Tarsal counts: 6, 8–9, 6, 6. Females (n = 5): Measurements: DSL: 4.61–4.75; DSW: 4.20–4.55; LI: 8.56–9.45; LII: 13.93–15.97; LIII: 10.25–11.28; LIV: 14.80–15.52. Pedipalps: Pedipalp tibial setation: mesal IiIi/IIi, lateral IiIi; tarsal setation: mesal IIi, lateral IiIi/Iiii. Legs: Femur IV proventral row of tubercles slightly enlarged or not. Patella IV with or without prodorsal apical projection. Tarsal counts: 5–6, 8–9, 6, 6.

‘Crassi-’ is a Greek adjective, meaning thick, and ‘tibia’ is the Latin noun meaning the podomere; combined to ‘crassitibia’ (nom. sing. fem.) in reference to the diagnostic male tibia IV of the species.

Brejos de Altitude humid montane forests of Ibiapaba mountains and Caatinga shrublands and dry forests of northern interior Ceará.

BRAZIL, Rio Grande do Norte: Portalegre, Cachoeira do Pinga.

Holotype. BRAZIL. Rio Grande do Norte: Portalegre, Cachoeira do Pinga, encosta da serra, elev. 431 m, 23–24.iv.2014, DaSilva M.B. and Saraiva N.E.V., ♂ (UFPB OP-481). Paratypes. same as for holotype, 3 ♂ (UFPB OP-482), 1 ♂ (UFPB OP-728) [dissected], 2 ♀ (UFPB OP-729) [female description], 1 ♂ (MZUSP 76647), 1 ♂ (MNRJ 60617).

Distinguished from other species by the presence of a pair of slightly enlarged paramedian tubercles on scute area III (Fig. 9A, C), instead of spines; coxa IV bearing a short, conical, straight prodorsal apical apophysis and a long, curved, sickle-like retro-apical apophysis (Fig. 9D); femur IV retro-lateral face with three central elongated apophysis (Fig. 10A, D); and contains five pairs of MS C (Fig. 11). Most similar to S. bicuspidata sp. nov. by the slightly elongated trochanter IV; penis glans’ stylus with a rounded dorso-apical projection; similar MS A and MS B; and MS D pairs close MS C group.

Holotype MALE (UFPB OP-481; Figs 9, 10): Measurements: DSL: 4.43; DSW: 4.87; LI: 8.91; LII: 19.48; LIII: 12.24; LIV: 16.53. Dorsum (Fig. 9A, C, G): Prosoma with few tubercles. Front margin bearing a single small median tubercle. Ocularium twice as tall as the eye diameter, with a pair of divergent spines slightly projected anteriorly (Fig. 9C, G). Two pairs of ozopores of similar size, anterior one placed laterally, and posterior one rounded and more dorsal; lateral channel developed and followed by oblong sensorial pegs that increase in size posteriorly. Scute area I with tubercles placed in a row along the longitudinal groove and scute groove II, with a pair of slightly enlarged paramedian tubercles (Fig. 9A). Scute areas II–IV each with a transversal central row of enlarged tubercles increasing in size towards the middle, paramedian pair larger than the others. Scute areas III–IV with more conical tubercles compared to those on other areas. Lateral margin of DS with an external row of low, rounded tubercles slightly increasing in size posteriorly. Posterior margin of DS and free tergites I–III each with a row of conical tubercles. Anal operculum tuberculate. Venter: Coxa I tuberculate, median row of large ones, apical face with two prolateral and two retro-lateral tubercles. Coxae II–III with small tubercles, pro- and retro-apical tubercles reduced. Coxa IV tuberculate. Chelicerae: Segment I with ectodorsal row of three tubercles on posterior face of bulla. Fixed and movable fingers with five and three teeth, respectively. Pedipalps (Fig. 9E, F): Trochanter inflated dorsally, ventral face with two setiferous tubercles, mesal largest. Femur with a basal setiferous tubercle, an external lateral row of 2–3 smaller setiferous tubercles on the middle 1/3 (Fig. 9E), and a mesal apical seta. Tibial setation: mesal IiIi, lateral IiIii. Tarsal setation: mesal IIi, lateral IiIi/Iii. Legs (Figs 9D, 10): Coxae I–III each with one prodorsal and one retro-dorsal apophysis; coxa II retro-dorsal apophysis trifid and fused at the apex with coxa III prodorsal apophysis. Coxa IV large, covered by more acuminate tubercles than those of the rest of the body (Fig. 9A); with an obliquely inserted, conical prodorsal apical apophysis that is slightly curved downwards (Fig. 9C, D); and a sickle-shaped retro-apical apophysis, twice the size of proapical apophysis, curved out and downwards (Figs 9D, 10D). Trochanter III with a reduced retro-apical projection. Trochanter IV long, with pro- and a retro-lateral central blunt apophyses, the prolateral one slightly acuminated (Fig. 10A); apical portion with a long, conical retro-apical apophysis and a small proapical round tubercle (Fig. 10). Femur III slightly curved retrolaterally in dorsal view, with a slightly acuminated retro-apical dorsal tubercle; ventral surface with two rows of tubercles increasing in size subapically, prolateral row larger. Femur IV slightly curved inwards at the base, medio-dorsal row of tubercles twice the height of those covering the podomere, almost on the entire femur (except the sub-apical surface), basal and sub-apical tubercles more conspicuous (Fig. 10A, B, D); a pair of long, curved dorso-apical paralateral spines, retro-lateral spine largest and strongly curved upwards (Fig. 10A, D); retro-lateral face with three long, conical, central apophyses (Fig. 10A, D); ventral surface with a prolateral row of large tubercles alternating with small ones on apical half, and a retro-lateral row with round tubercles (Fig. 10B, C); ventro-apical portion with two paralateral large straight spiniform apophyses, proapical largest (Fig. 10C). Patella IV tuberculate, ventral surface covered by enlarged, acuminated tubercles, with a large proapical spine (twice the size of the surrounding ones) (Fig. 10B). Tibia IV with two ventral rows of tubercles increasing in size apically, pointing posteriorly (Fig. 10B, D). Metatarsus I slightly swollen. Tarsal counts: 6, 9, 6, 6. Penis (UFPB OP-728; Fig. 11): VP apex and lateral margins slightly concave. MS A set showing asymmetry between right (4) and left (3) sides (Fig. 10A, B); MS B almost undistinguishable from MS A but inserted more basally. Five pairs of conical, curved MS C, decreasing in size apically; two pairs of MS D close to MS C5, but slightly smaller and placed more dorsally; two pairs of small MS E, ventral to the MS C set (Fig. 10B). Coloration: General coloration predominantly moderate yellow (87). DS scute areas, prosoma and margins, free tergites I–III moderate olive brown (95) granulation.

FEMALE (UFPB OP-729; Figs 6B, 12D) Measurements: DSL: 4.13; DSW: 4.32; LI: 8.83; LII: 16.57; LIII: 10.93; LIV: 14.40. Dorsum: Less armed compared to male. DS narrower. Lateral margin of DS with an external row of tubercles, posterior tubercles slightly larger than in male. Pedipalps: Pedipalp tibial setation: lateral IiIi. Legs: Coxa IV narrower and shorter than in male, with a small prolateral apophysis, and a conical, straight retro-apical apophysis of same size. Trochanter IV narrower and shorter than in male, the prolateral basal apophysis as a large, blunt tubercle, and a small retro-lateral apophysis. Femur III lacking a proventral row of enlarged tubercles, reduced retro-dorsal apical tubercle. Femur IV unarmed, covered by small conical tubercles that slightly increase in size apically, with a pair of small dorso-apical spines of similar size. Patella–tibia IV unarmed. Tarsal counts: 6, 9–10, 6, 6.

Males (n = 6): Measurements: DSL 4.43–4.66; DSW 4.84–5.06; LI 8.91–10.21; LII 17.52–19.48; LIII 12.24–13.08; LIV 16.19–17.51. Dorsum: Front margin with or without tubercles. Ocularium with parallel or diverging spines, inclination of these ranging from near-horizontal to vertical. Pedipalps: Femur external lateral row with 2–3 sub-apical tubercles. Legs: Femur IV dorso-apical spines with similar size to retro-lateral largest and strongly curved upwards; retro-lateral central conical apophyses sometimes with an additional apophysis of half the height of the remaining apophyses in the middle; apical half of proventral row bearing small tubercles alternated with large ones. Tarsal counts: 6, 10–12, 6, 4–6. Females (n = 2): Measurements: DSL 4.13–4.27; DSW 4.32–4.46; LI 8.83–8.84; LII 16.16–16.57; LIII 10.93–11.62; LIV 10.09. Pedipalps: Pedipalp tibial setation: mesal IiIi, lateral IiIi. Tarsal setation: mesal IIi, lateral IiIi/IiIii. Legs: Tarsal counts: 6, 9–10, 6, 6.

‘Falcata’ is a Latin adjective (adj. nom. fem.) that means sickle-shaped and refers to the diagnostic retro-apical apophysis of male coxa IV of the species.

Known only from type locality. Caatinga dry forest of tableland slopes of interior Rio Grande do Norte.