2.1. Taxon sampling

2.2. DNA extraction, PCR amplification and sequencing

2.3. Phylogenetic analysis

2.4. Morphological study

3.1. The subgenus Cryobius is recovered as a monophyletic group within Pterostichus

3.2. The Pyreneo-Cantabrian Cryobius

3.3. P. pumilio

3.4. The P. abaxoides- and pusillus groups

3.5. The P. cantabricus group

3.6. Cryobius from the Alps

3.7. The Nearctic and Eastern Oriental Cryobius

3.8. P. cf. anatolicus

4.1. The subgenus Cryobius

4.2. Haptoderus and Pyreneorites

4.3. The Pyreneo-Cantabrian Cryobius

4.4. The Pyrenean Cryobius

4.5. The Cantabrian clade

4.6. The case of the widespread species P. pumilio

4.7. The Eastern Cryobius

4.8. The Nearctic Cryobius