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  • ContentsContents
  • Article InfoArticle Info
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  • TaxaTaxa
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  • Article title
  • Abstract
  • Keywords
  • 1. Introduction
    • 1.1. The earliest beetles and their systematic treatment
    • 1.2. Reflectance Transformation Imaging
  • 2. Material and methods
    • 2.1. Material
    • 2.2. Imaging
    • 2.3. RTI dome
    • 2.4. RTI processing
    • 2.5. Image processing
    • 2.6. Graphic design
    • 2.7. Data availability
  • 3. Results
    • Coleopsis
    • Coleopsis archaica
  • 4. Discussion
    • 4.1 New interpretation of morphological features
    • 4.2. Systematic interpretation
    • 4.3. Coleopteran autapomorphic character states documented in the fossil (features shared with other extinct and extant beetles)
    • 4.4. Plesiomorphic character states of the fossil (features not shared with extant beetles)
    • 4.5. Possible affinities within Coleoptera
    • 4.6. RTI technique
  • 5. Conclusions
  • 6. Acknowledgements
  • 7. References
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