Arthropod Systematics & Phylogeny 72(3): 223-236, doi: 10.3897/asp.72.e31787
Description of a new species of Glaresidae (Coleoptera: Scarabaeoidea) from the Jehol Biota of China with a geometric morphometric evaluation
expand article infoMing Bai, Rolf Beutel‡, Wangang Liu, Sha Li, Mengna Zhang, Yuanyuan Lu, Keqing Song, Dong Ren§, Xing-Ke Yang|
‡ University of Jena, Jena, Germany§ Capital of Normal University, Beijing, China| Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Corresponding author: Dong Ren ( rendong@cnu.edu.cn )

Corresponding author: Xing-Ke Yang ( yangxk@ioz.ac.cn )

© Ming Bai, Rolf Beutel, Wangang Liu, Sha Li, Mengna Zhang, Yuanyuan Lu, Keqing Song, Dong Ren, Xing-Ke Yang.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Open Access
Abstract
Glaresis tridentata Bai, Beutel & Ren sp. nov. (Scarabaeoidea: Glaresidae) from the Yixian Formation of western Liaoning Province, NE China is described and illustrated. A geometric morphometric analysis of three character systems (head, metatibia and aedeagus) was carried out including 136 specimens from 44 species of three genera of extant and extinct Glaresidae. The variation in these features between Glaresis tridentata Bai, Beutel & Ren sp. nov. and the other species was estimated based on Principal Component Analysis (PCA), Canonical Variate Analysis (CVA), Discriminant function analysis (DFA) and Minimum Spanning Tree (MST) approaches. The results suggest that Glaresidae, likely the sister group of all other extant Scarabaeoidea, had been established in the Early Cretaceous and contained larger species than today. The assignement of the new species to the genus Glaresis is supported. The validity of the genera Lithoglaresis and Cretoglaresis is questionable.
Keywords
Fossil, Glaresidae, Yixian Formation, new species, geometric morphometrics.1. IntroductionGlaresidae is a small and uniform family of Scarabaeoideaoccurring on all continents except for Australia andAntarctica. About