Arthropod Systematics & Phylogeny 73(2): 259-279, doi: 10.3897/asp.73.e31800
The higher phylogeny of Leptophlebiidae (Insecta: Ephemeroptera), with description of a new species of Calliarcys Eaton, 1881
expand article infoRoman J. J. Godunko§|, Pavel Sroka, Tomáš Soldán, Jindřiška Bojková
‡ State Museum of Natural History, National Academy of Sciences of Ukraine, Lviv, Ukraine§ Biology Centre, Czech Academy of Sciences, Institute of Entomology, České Budějovice, Czech Republic| University of Łódź, Łódź, Poland
Open Access
A detailed morphological description of the adults and larvae of Calliarcys van sp.n., the second representative of the genus Calliarcys Eaton, 1881 is provided, including a differential diagnosis with regard to Calliarcys humilis Eaton, 1881. C. van sp.n. is distinguished in the adult mainly by the colouration of the upper part of male compound eyes, shape of costal process of hind wings, and genitalia; in larvae it is distinguished by the shape of hypopharynx and labium; and by comparatively densely arranged unbranched longitudinal ridges on the exochorion of the fusiform eggs. The generic diagnosis for Calliarcys is re-evaluated and specified. The genus had already previously been proposed to constitute the subfamily Calliarcyinae within Leptophlebiidae, a concept which has not been generally accepted. Now this classification is tested by phylogenetic analysis, for the first time using a set of 20 morphological characters. Ambiguous characters from previous studies are discussed. Leptophlebiinae is confirmed as sister group to all remaining Leptophlebiidae. Calliarcys is revealed as sistergroup of Habrophlebiinae + Atalophlebiinae s.l., thus justifying its classification as a subfamily Calliarcyinae.
Calliarcyinae, systematics, phylogeny, Asia Minor.